Uplistsikhe's old main street winds up to the right after you've passed through the main gate, with several important cave structures either side of it.
Down to the right is the large pre-Christian Temple of Makvliani, with an inner recess behind an arched portico. A little further up on the left is the big hall known as Tamaris Darbazi (Hall of Queen Tamar), where an ancient stone seat sits behind two columns cut from the rock, and the stone ceiling is carved to look like wooden beams. This was almost certainly a pre-Christian temple. To its left is an area with stone niches, thought to have been a pharmacy or dovecote.
A large cave building to the right of the Tamaris Darbazi was probably a sun temple used for animal sacrifices, later converted into a Christian basilica.